The El Al airliner made the three-hour trip, carrying a delegation of Israeli and US officials.
The flight was allowed to cross Saudi Arabian airspace, normally blocked to Israeli air traffic.
The UAE became only the third Arab country in the Middle East to recognise Israel since its founding in 1948.
On Saturday the UAE repealed a law boycotting Israel which had been in place since 1972, and earlier this month the two countries opened direct telephone services for the first time.
The agreement to normalise relations - brokered by the US - was made public in a surprise announcement on 13 August.
Flight LY971 - numbered to represent the UAE's international dialling code - carried delegates including Donald Trump's son-in-law and senior adviser Jared Kushner and Israel's National Security Adviser Meir Ben-Shabbat.
Mr Kushner led secret talks which resulted in the Israel-UAE agreement.
The joint teams will meet Emirati representatives to develop areas of co-operation between Israel and the UAE. The return flight will be numbered LY972, after Israel's international dialling code.
Monday's three-hour flight has taken more than 70 years to make, and it marks a new turning point in relations between Israel and the Arab world.
There are big prizes for all three players: Israel's historic need to boost regional recognition of the Jewish state (could Saudi Arabia one day do so too?); the Emiratis' glittering finance hubs can benefit from open links with the region's security and cyber superpower; while a US president under pressure at home gets to tout his role as peacemaker in the Middle East.
These are truly significant achievements and further shift the dynamics in a deeply polarised region. But the deal is striking for another reason - it leaves the Palestinians feeling as sidelined as ever.
They believe it breaks years of Arab solidarity - and leverage - against Israel's occupation of land they want for a future state; while ordinary Palestinians feel more and more hemmed in as Israeli settler numbers grow.
They see not only betrayal, but a blind eye being turned by the Emiratis to their reality on the ground.
While it was welcomed by much of the international community, the UAE's recognition of Israel without the precondition of the creation of a Palestinian state was denounced by the Palestinians as a betrayal of their cause.
In return for official relations with the UAE, Mr Netanyahu agreed to suspend controversial plans to annex parts of the occupied West Bank - land claimed by the Palestinians for a future state of their own.
Before the UAE, Egypt and Jordan were the only other Arab countries in the Middle East to officially recognise Israel, after signing peace treaties in 1978 and 1994 respectively.
Mauritania, a member of the Arab League in north-west Africa, established diplomatic relations with Israel in 1999 but severed ties in 2010.
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